2007 – 2013        PhD Ecology and Evolutionary Biology, University of Arizona

2002 – 2006        BA Biology and Chemistry, University of Pennsylvania

Research & Professional Experience

2018-      Assistant Professor, Department of Biology, Virginia Commonwealth University

2014-17    Assistant Professor, Department of Biology, University of Puerto Rico

2013     NSF Postdoctoral Research Fellow, University of California Davis

2012     Adjunct Faculty, Pima Community College Desert Vista

2012     Graduate Teaching Associate, Introduction to Global Change

2012     Graduate Teaching Associate, Evolutionary Biology

2009     Grant Writer, Environmental Education Exchange

2008     Research Associate, Conservation International

2007     Laboratory Technician, Helliker Lab, University of Pennsylvania

2002-06      Research Technician, Janzen Lab, University of Pennsylvania


Wieczynski D, Boyle B, Buzzard V, Duran S, Henderson A, Hulshof CM, Kerkhoff A, McCarthy M, Michaletz S, Swenson S, Asner G, Bentley L, Enquist B, Savage V. 2019. Climate shapes and shifts functional biodiversity in forests worldwide. Nature 116: 587-592.

Derroire G, Powers J, Hulshof CM, Varela L, Healy J. 2018. Contrasting patterns of leaf trait variation among and within species during tropical dry forest succession in Costa Rica. Nature Scientific Reports 8: 285.

Agosta SJ, Hulshof CM, Staats E. 2017. Herbivore performance, climate, and leaf traits in regenerating tropical dry forests. Journal of Animal Ecology 86: 590-604.

Allen K, Dupuy JM, Gei MG, Hulshof CM, Medvigy D, Pizano C, Salgado-Negret B, Smith CM, Trierweiler A, Van Bloem SJ, Waring BG, Xu X, Powers JS. 2017. Will seasonally dry tropical forests be sensitive or resistant to future changes in rainfall regimes? Special Issue Environmental Research Letters 12: 023001s. Featured in Highlights of 2017 collection from Environmental Research Letters.

Buzzard V, Hulshof CM, Violle C, Enquist BJ. 2016. Regrowing a tropical dry forest: Functional trait diversity during secondary succession. Functional Ecology 30: 1006-1013.

Hulshof CM, Swensons N, Weiser M. 2015. Tree height-diameter allometry across the United States. Ecology and Evolution 5: 1193-1204

Hulshof CM, Violle C, Spasojevic M, McGill B, Damschen E, Harrison S, Enquist B. 2013. The relative importance of intra- and interspecific trait variation in the maintenance of plant species diversity. Journal of Vegetation Science 24: 921-931.

Hulshof CM, Martinez-Yrizar A, Burquez A, Boyle B, Enquist B. 2013. Functional plant trait variation in tropical dry forests: A review and synthesis. In Tropical Dry forests of the Americas: Ecology, Conservation, and Management (eds. J. Powers and A. Sanchez), CRC Press, New York. pp 129-140.

Violle C, Enquist B, McGill B, Jiang L, Albert C, Hulshof CM, Jung V, Messier J. 2012. The return of the variance: intraspecific variability in community ecology. TREE 27: 244-252.

Hulshof CM, Swenson NG, Stegen JJ, Enquist CF, Enquist BJ. 2012. Interannual variability of growth and reproduction in the tropical tree Bursera simaruba – The role of allometry and resource variability. Ecology 93:180–190.

Hulshof CM, Swenson NG. 2010. Variation in leaf functional trait values within and across individuals and species: An example from a Costa Rican dry forest. Functional Ecology 24: 217 – 223.

In Press, Accepted, or In Review

Grants, Awards, & Fellowships

2017     National Science Foundation EPSCoR Research Fellowship (declined due to move)

2016     National Science Foundation Macrosystems Biology Early Career Award

2016     Puerto Rico Science, Technology, and Research Trust Small Research Grant

2016     NASA EPSCoR IDEAS Small Grant Award

2015     U.S. Fish and Wildlife Cooperative Research Grant

2015     Association of Tropical Biology and Conservation Travel Award

2014     Carl Storm Underrepresented Minority Fellowship

2012     National Science Foundation Postdoctoral Fellowship in Biology

2012     Ecological Society of America Student Travel Award

2012     Institute of the Environment Graduate Student Travel Award

2011     Graduate and Professionals Student Council Travel Award

2010     Philanthropic Educational Organization Scholar Award Nominee

2010     Institute of the Environment Dissertation Improvement Grant

2009     National Science Foundation Graduate Research Fellowship

2009     Ford Foundation Diversity Fellowship Doctoral Program

2009     Tinker Summer Field Research Grant

2008     NSF/OTS Graduate Student International Fellowship

2008     NSF-Amazon PIRE Travel Grant, Tapajós National Forest, Brazil

2007     Graduate Diversity Fellowship, University of Arizona

2006     Nassau Research Fund, University of Pennsylvania

2006     Binns-Williams Research Fund, University of Pennsylvania

2005     NSF-Research Experience for Undergraduates (REU) Fellowship

Courses Taught: Data Science for Biologists; General Biology; Principles of Ecology; Quantitative Analyses in R; Modeling in Population Ecology

Professional Service

Peer Reviewer: American Naturalist, Annals of Botany, Biotropica, Ecology, Functional Ecology, Global Ecology and Biodiversity, Journal of Animal Ecology, Journal of Biogeography, Journal of Ecology, Oecologia, Oikos, PlosONE, Proceedings of the Royal Society B, among others

Professional memberships: Association of Tropical Biology and Conservation; Ecological Society of America; Faculty advisor for UPRM ESA-SEEDS student chapter (2014-2017)